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SAN JOSE, Calif. — Tristan Begg was an anthropology student at UC Santa Cruz and a Beethoven fanatic when he volunteered as a docent at San Jose State University’s Beethoven center in the summer of 2009. He would pull out the drawer holding a lock of hair and tell visitors, “This is real” and that it once was on the head of the greatest composer who ever lived, the one whose music changed Begg’s life when he heard the first notes of Moonlight Sonata on Christmas morning at age 17. “It was instantaneous. I was astounded....